Marine invertebrate surveys

Collaborators: Wilma Blom (Auckland War Memorial Museum); Fred Brook (Whangarei); Brett Stephenson (Whakatane)

This programme is jointly co-ordinated by Bruce Hayward and Margaret Morley. The goal of this "hobby" research programme is to document the biodiversity and biogeography,and map the ecological distribution of the intertidal biota and inner shelf subtidal soft sediment benthos around the coast of northern New Zealand. All phyla are covered with emphasis on the molluscs, seaweeds, and echinoderms.

Study areas (to date):
a. New Plymouth (proposed Nga Motu Marine Reserve). Published.
b. northern Taranaki (New Plymouth to Awakino). Published.
c. Kawhia Harbour. Published.
d. Raglan Harbour and Karioi coast. Published.
e. Waitakeres (Muriwai to Big Muddy Ck). Published.
f. Kawerua (Waipoua coast). Published.
g. Whangape Harbour. Published.
h. Ahipara and Herekino Harbour. Published.
i. Parengarenga Harbour. Published.
j. Cavalli Islands (subtidal benthos). Published.
k. Whananaki Harbour and coast. Published.
l. North-east Great Barrier (Rakitu, Rangiwhakaea Bay). Published.
m. West Great Barrier (Port Fitzroy, Port Abercrombie). Write-up phase.
n. Broken Islands subtidal benthos (west Great Barrier). Published.
o. Great Barrier mollusca. Published, 2009.
p. Middle and Upper Waitemata Harbour. (map) Published.
q. Waitemata Harbour (subtidal benthos). Part published.
r. Tamaki Estuary (mapping). Published, 2008.
s. Hauraki Gulf intertidal. In progress 2003-2012.
t. Te Matuku Bay Marine Reserve (Waiheke). Published.
u. North-east Coromandel (Whangapoua). Published 2014.
v. Cuvier Island. Published.
w. Mahia Peninsula. Published.
x. Bay of Islands subtidal. Part published.
y. Subantarctic Campbell and Auckland Islands. Published.
z. Introduced tunicates Pyura praeputialis and Styela clava to NZ. Published, 2009

aa. Spread of the blue mussel. Published, 2010.

bb. Intertidal Nudibranchs. Submitted 2015.

cc. Intertidal micromolluscs. Write up phase, 2015.



  • Hayward, B.W., Morley, M.S., Stephenson, A.B., Grenfell, H.R., Hayward, G.C., 2014. Intertidal shallow and subtidal biota, northeast Coromandel Peninsula. Poirieria 38, 2-12.
  • Morley, M.S., Hayward, B.W., 2014. Biodiversity and distribution of Ostracoda, Foraminifera and Micromollusca of Matai Bay, Northland. Records of Auckland Museum 49, 55-80.
  • Hayward, B.W., Morley, M.S., Grenfell, H.R. 2012: Marine biota of the Whananaki coast, east Northland, New Zealand. Records of Auckland Museum 48: 77-96.
  • Morley, M.S., Hayward, B.W., and Hipkins, K., 2011. Manukau Harbour Mollusc survey, 1952-1963. Poirieria 36: 24-36.
  • Hayward, B.W., Morley, M.S., 2010: Hinea brasiliana alive and well at North Cape. Poirieria 35, 15-18.
  • Morley, M.S., Hayward, B.W., 2010: The blue mussel, Mytilus galloprovincialis planulatus, expands its range around Auckland. Poirieria 35, 25-30.
  • Beu, A.G., Hayward, B.W., 2010: Mollusca dredged from Port Pegasus, Stewart Island. Poirieria 35, 7-10.
  • Morley, M.S., Hayward, B.W., 2009: Marine Mollusca of Great Barrier Island, New Zealand. Records of Auckland Museum 46, 15-51.
  • Hayward, B.W., Morley, M.S., 2009: Introduction to New Zealand of two sea squirts (Tunicata, Ascidacea) and their subsequent dispersal. Records of Auckland Museum 46, 5-14.
  • Hayward, B.W., 2009, Mass mating of the mud snail, Amphibola crenata: Poirieria, v. 34, p. 23.
  • Hayward, B.W., and Morley, M.S., 2008, Intertidal life of the Tamaki Estuary and its entrance, Auckland. Auckland Regional Council Technical Publication.373, 71 pp.
  • Morley, M.S.; Hayward, B.W.; Raven, J.L.; Foreman, G.A.; Grenfell, H.R., 2006. Intertidal and shallow subtidal biota of Mahia Peninsula, Hawke Bay. Records of Auckland Institute and Museum 43: 11-41.
  • Hayward, B.W.; Morley, M.S., 2005. Zonation and biogeography of the intertidal biota of Subantarctic Campbell and Auckland Islands, New Zealand. Records of Auckland Institute and Museum 42: 7-33.
  • Hayward, B.W. (Scientific editor), 2004. Morton, J.E. Seashore Ecology of New Zealand and the Pacific. David Bateman. Auckland. 504 p.
  • Hayward, B.W.; Morley, M.S.; Grenfell, H.R.; Carter, R.; Hayward, G.C.; Blom, W.M.; Rogan, D., 2004. Intertidal biota and washup at Ahipara and Herekino, Northland west coast. Poirieria 30: 13-25.
  • Hayward, B.W.; Morley, M.S., 2003. Intertidal and shallow subtidal biota of Cuvier Island. Poirieria 29: 16-23.
  • Hayward, B.W.; Morley, M.S., 2004. Intertidal life around the coast of the Waitakere Ranges, Auckland. Auckland Regional Council Working Report No. 111. 102 p.
  • Hayward, B.W.; Morley, M.S., 2002. Intertidal biota of the proposed Nga Motu Marine Reserve, New Plymouth. Poirieria 28: 1-11.
  • Hayward, B.W.; Morley, M.S., 2002. Marine biota of Raglan, Waikato West Coast. Poirieria 28: 1-16.
  • Hayward, B.W.; Stephenson, A.B.; Morley, M.S.; Blom, W.M.; Grenfell, H.R.; Brook, F.J.; Riley, J.J.; Thompson, F.; Hayward, J.J., 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of Auckland Museum 37: 45-80.
  • Hayward, B.W.; Morley, M.S.; Hayward, J.J.; Stephenson, A.B.; Blom, W.M.; Hayward, K.A.; Grenfell, H.R., 1999. Monitoring studies of the benthic ecology of Waitemata Harbour, New Zealand. Records of the Auckland Museum 36: 95-117.
  • Hayward, B.W.; Morley, M.S.; Stephenson, A.B.; Blom, W.; Grenfell, H.R.; Prasad, R., 1999. Marine biota of the north Taranaki coast, New Zealand. Tane 37: 171-199.
  • Hayward, B.W.; Morley, M.S.; Stephenson, A.B.; Blom, W.M.; Grenfell, H.R.; Prasad, R.; Rogan, D.; Thompson, F.; Cheetham, J.; Webb, M., 1999. Intertidal and subtidal biota and habitats of the central Waitemata Harbour. Auckland Regional Council Technical Publication No. 127. 40 p.
  • Hayward, C.M.; Hayward, B.W., 1999. Human impact on Orakei Basin, Auckland. Tane 37: 137-152.
  • Morley, M.S.; Hayward, B.W., 1999. Inner shelf mollusca of the Bay of Islands, New Zealand, and their depth distribution. Records of the Auckland Museum 36: 119-138.
  • Hayward, B.W., 1997. Introduced marine organisms in New Zealand and their impact in the Waitemata Harbour, Auckland. Tane 36: 197-223.
  • Hayward, B.W.; Stephenson, A.B.; Morley, M.S.; Riley, J.; Grenfell, H.R., 1997. Faunal changes in Waitemata Harbour sediments, 1930s-1990s. Journal of the Royal Society of New Zealand 27(1): 1-20.
  • Hayward, B.W.; Stephenson, A.B.; Morley, M.S.; Smith, N.; Thompson, F.; Blom, W.; Stace, G.; Riley, J.L.; Prasad, R.; Reid, C., 1997. Intertidal biota of Te Matuku Bay, Waiheke Island, Auckland. Tane 36: 67-84.
  • Hayward, B.W.; Grace, R.V.; McCallum, J., 1986. Soft bottom macrobenthos and sediments off the Broken Islands, northern New Zealand. Tane 31: 85-103